A decade has passed since the Ant Queen's apocalyptic war with the humans. The queen's conscripts in the war, a race of uplifted animals, now make their way in the world they helped destroy. And after years of bloodshed, the survivors finally have a fleeting chance to build a lasting peace. But peace always comes with a price. The holy city of Hosanna - where animals and humans form a joint government - finds itself surrounded by wolves determined to retake the land. A powerful matriarch has united the rival wolf packs, using a terrible power harnessed from the Queen herself.