Matt Reddy and his crew fight alongside the felinoid Lemurians and Imperial allies to keep the reptilian Grik, a race growing in supremacy, from reconquering the Lemurians' ancestral home on Madagascar. Exhausted, far from reinforcements and outnumbered, the odds are greater than ever. As for the fate of the Americas, Don Hernan and the evil Dominion have gathered to annihilate the forces behind the walls of Fort Defiance as a shadowy power with an agenda all its own rises with chilling resolve.